SDSU EB Vertebrate Collections

Welcome to the Research & Publications Page

Our collections are used for research as well as teaching. Here is a partial list of research theses and publications that have used specimens in our collection. To view a list of publications from 2009 and earlier please click here.

Bird Collection


Barker, F. K., K. J. Burns, J. Klicka, S. M. Lanyon, and I. J. Lovette. 2013. Going to extremes: Contrasting rates of diversification in a recent radiation of New World passerine birds. Systematic Biology 62: 298-320.

Maley, J. M., R. T. Brumfield. 2013. Mitochondrial and Next-Generation Sequence Data used to Infer Phylogenetic Relationships and Species Limits in the Clapper/ King Rail Complex. The Condor 115: 316-329.  


Burns, K. J. and A. J. Shultz. 2012. Widespread cryptic dichromatism and ultravioloet reflectance in the largest radiation of Neotropical songbirds: implications of accounting for avian vision in the study of plumage evolution. The Auk 129: 211-221.

Moyle, R. G., M. J. Anderson, C. H. Oliveros, F. D. Steinheimer, and S. Reddy. 2012. Phylogeny and Biogeography of the Core Babblers (Aves: Timaliidae) Systematic Biolpgy 61: 631-651.


Caballero, I. C. and M. V. Ashley. 2011. Genetic analysis of the endemic island loggerhead shrike, Lanius ludovicianus anthonyi. Conservation Genetics 12: 1485-1493.

Lovette, I. J., B. S. Arbogast, B. Curry, R. M. Zink, C. A. Botero, J. P. Sullivan, A. L. Talaba, R. B. Harris, D. R. Rubenstein, R. E. Ricklefs, and E. Bermingham. 2011. Phylogenetic relationships of the mockingbirds and thrashers (Aves: Mimidae). Molecular Phylogenetics and Evolution 63: 219-229.

Shultz, A. J. 2011. Using an avian visual model to assess the evolution of sexual dichromatism in tanagers. MS Thesis, San Diego State University.  

Herbarium Collection


Benet-Pierce, N. and M. G. Simpson. 2014. The taxonomy of Chenopodium desiccatum and C. nitens, sp. nov. Journal of the Torrey Botanical Society 141:161-172. (pdf)

Rebman, J. P. and M. G. Simpson. 2014. Checklist of the Vascular Plants of San Diego County, 5th edition. San Diego Natural History Museum Publications.

Ripma, L.*, M. G. Simpson, and K. Hasenstab-Lehman*. 2014. Geneious! Simplified genome skimming methods for phylogenetic systematic studies: A case study in Oreocarya (Boraginaceae). Applications in Plant Sciences doi: 10.3732/apps.1400062. (pdf)

Simpson, M. G., R. Dowdy*, J. P. Rebman, R. B. Kelley, and L. M. Simpson. 2014. The recognition of two species in Eremocarya (Boraginaceae): Evidence from fornix bodies, nutlets, corolla size, and biogeography. Madroño 61: 259–275. (pdf)


Guilliams, C. M.*, B. A. Veno, M. G. Simpson, and R. B. Kelley. 2013. Pectocarya anisocarpa, a new species of Boraginaceae, and a revised key for the genus in western North America. Aliso 31, 1-13. (pdf)

Silveira, M.* and Simpson, M. G. 2013. Phylogenetic systematics of the mesa mints: Pogogyne (Lamiaceae). Systematic Botany 38: 782-794. (pdf)

Simpson, M. G. and J. P. Rebman. 2013. A new species of Cryptantha (Boraginaceae) from the Sierra de San Pedro Mártir, Baja California, Mexico. Madroño 60(1): 35-45. (pdf)

Simpson, M. G., J. P. Rebman, K. E. Hasenstab-Lehman, C. M. Guilliams, and P. McConnell. 2013. Cryptantha wigginsii (Boraginaceae): A presumed extinct species rediscovered. Madroño 60(1): 24-34. (pdf)

Yost, J. M., M. Bontrager, S. McCabe, D. Burton*, M. G. Simpson, K. M. Kay, and M. Ritter. 2013. Phylogenetic relationships and evolution in Dudleya (Crassulaceae). Systematic Botany 38, 1096-1104. (pdf)


Dodero, M. W.* and M. G. Simpson. 2012. Dudleya crassifolia (Crassulaceae), a new species from northern Baja California, Mexico. Madroño 59: 223-229. (pdf)

Hasenstab-Lehman, Kristen E.* and M. G. Simpson. 2012. Cat’s Eyes and Popcorn Flowers: Phylogenetic Systematics of the Genus Cryptantha s.l. (Boraginaceae). Systematic Botany 37: 738–757. (pdf)

Kelley, R. B., M. G. Simpson, and K. E. Hasenstab. 2012. Cryptantha. Pp. 455-468 in B. G. Baldwin, D. H. Goldman, D. J. Keil, R. Patterson, T. J. Rosatti, and D. H. Wilken (eds.), The Jepson Manual: Vascular Plants of California. Second Edition. University of California Press, Berkeley, Los Angeles, London.

Silveira, M., M. G. Simpson, and J. D. Jokerst. 2012. Pogogyne. Pp. 850-851 in B. G. Baldwin, D. H. Goldman, D. J. Keil, R. Patterson, T. J. Rosatti, and D. H. Wilken (eds.), The Jepson Manual: Vascular Plants of California. Second Edition. University of California Press, Berkeley, Los Angeles, London.  


Guilliams, C. M.*, Simpson, M. G., and J. P. Rebman. 2011. Calyptridium parryi var. martirense (Montiaceae), a new taxon endemic to the Sierra San Pedro Mártir, Baja California, Mexico. Madroño 58(4): 258-266. (pdf)


Benet-Pierce, N. and Michael G. Simpson. 2010. Chenopodium littoreum (Chenopodiaceae), a new goosefoot from dunes of south-central coastal California. Madroño 57: 64-72. (pdf)

Simpson, Michael G., C. M. Guilliams*, and M. Silveira*. 2010. Taxonomy of Calyptridium parryi (Montiaceae). Madroño 57: 145-160. (pdf)

Herpetology Collection


Torres-Carvajal, O., R. Etheridge and K. de Queroz. 2011. A systematic revision of neotropical lizards in the calde Hoplocercinae (Squamata: Iguania). Zootaxa 2752: 1-44.


Online Publications (2010-2011)

Mammal Collection


Berta, A., E. G. Ekdale, N. Zellmer, T. A. Deméré, S. Kienle, M. Smallcomb. 2015. Eye, nose, hair, and throat: external anatomy of the head of Eschrichtius robustus (Cetacea, Mysticeti). Anatomical Record 298:648-659.

Ekdale, E. G., T. A. Deméré, and A. Berta. 2015. Vascularization of the gray whale palate (Cetacea, Mysticeti, Eschrichtius robustus): soft tissue evidence for an alveolar source of blood to baleen. Anatomical Record 298:691-702.

El Adli, J. E. and T. A. Deméré. 2015. On the anatomy of the temporomandibular joint and the muscles that act upon it: observations on the gray whale, Eschrichtius robustus. Anatomical Record 298:680-690.


Ekdale, E. G., A. Berta, and T. A. Deméré. 2011. The comparative osteology of the petrotympanic complex (ear region) of extant baleen whales (Cetacea: Mysticeti). PLoS ONE 6(6):e21311. doi: 10.1371/journal.pone.0021311. Published in PLoS ONE.

Walsh, B. and *A. Berta.* 2011. Occipital ossification of balaenopteroid mysticetes. *The* *Anatomical Record*294:391-398. (pdf)


Johnston, C., T. Deméré, *A. Berta*, J. St. Leger and J. Yonas. 2010. Observations on the musculoskeletal anatomy of the head of a neonate gray whale (*Eschrichtius robustus*). *Marine Mammal Science, *26 (1): 186-194. 

Terrestrial Arthropod Collection


M Hedin. 2015. High stakes species delimitation in eyeless cave spiders (Cicurina, Dictynidae, Araneae) from central Texas. Molecular Ecology 24, 346-361. 


Derkarabetian S, M Hedin. 2014. Integrative taxonomy and species delimitation in harvestmen: A revision of the western North American genus Sclerobunus (Opiliones: Laniatores: Travunioidea). PLoS ONE 9(8): e104982.


Hedin M, J Starrett & C Hayashi. 2013. Crossing the uncrossable: Novel trans-valley biogeographic patterns revealed in the genetic history of low dispersal mygalomorph spiders (Antrodiaetidae, Antrodiaetus) from California. Molecular Ecology 22, 508-526.

Richart CH and M Hedin. 2013. Three new species in the harvestmen genus Acuclavella (Opiliones, Dyspnoi, Ischyropsalidoidea), including description of male Acuclavella quattuor (Shear 1986). Zookeys 311, 19-68.

Satler JD, BC Carstens, M Hedin. 2013. Multilocus species delimitation in a complex of morphologically conserved trapdoor spiders (Mygalomorphae, Antrodiaetidae, Aliatypus). Systematic Biology 62, 805–823.

Schönhofer AL, M McCormack, N Tsurusaki, J Martens & M Hedin. 2013. Molecular phylogeny of the harvestmen genus Sabacon (Arachnida: Opiliones: Dyspnoi) reveals multiple Eocene-Oligocene intercontinental dispersal events in the Holarctic. Molecular Phylogenetics and Evolution 66, 303-315.  


Bond JE, BE Hendrixson, CA Hamilton & M Hedin. 2012. A reconsideration of the classification of the spider infraorder Mygalomorphae based on three nuclear genes and morphology (Arachnida: Araneae).PLoS ONE 7(6): e38753. doi:10.1371/journal.pone.0038753.

Burns M, M Hedin & JW Shultz. 2012. Molecular phylogeny of the leiobunine harvestmen of eastern North America (Opiliones: Sclerosomatidae: Leiobuninae). Molecular Phylogenetics and Evolution 63, 291-298.

Hedin M, J Starrett, S Akhter, AL Schönhofer & JW Shultz. 2012. Phylogenomic resolution of Paleozoic divergences in harvestmen (Arachnida, Opiliones) via analysis of next-generation transcriptome data. PLoS ONE 7(8): e42888. doi:10.1371/journal.pone.0042888

Hedin M, N Tsurusaki, R Macías-Ordóñez & JW Shultz.. 2012. Molecular systematics of sclerosomatid harvestmen (Opiliones, Phalangioidea, Sclerosomatidae): geography is better than taxonomy in predicting phylogeny. Molecular Phylogenetics and Evolution 62, 224-236.

Keith RM & M Hedin. 2012. Extreme mitochondrial population subdivision in southern Appalachian paleoendemic spiders (Araneae, Hypochilidae, Hypochilus), with implications for species delimitation. Journal of Arachnology 40, 167-181.


Derkarabetian S, J Ledford & M Hedin. 2011. Genetic diversification without obvious genitalic morphological divergence in harvestmen (Opiliones, Laniatores, Sclerobunus robustus) from montane sky islands of western North America. Molecular Phylogenetics and Evolution 61, 844-853.

Hedin M & D Carlson. 2011. A new trapdoor spider species from the southern Coast Ranges of California (Mygalomorphae, Antrodiaetidae, Aliatypus coylei, nov sp), including consideration of mitochondrial phylogeographic structuring. Zootaxa 2963, 55-68.

Satler J, J Starrett, C Hayashi & M Hedin. 2011. Inferring species trees from gene trees in a radiation of California trapdoor spiders (Araneae, Antrodiaetidae, Aliatypus). PLoS ONE 6(9): e25355. doi:10.1371/journal.pone.0025355.


Derkarabetian S, DB Steinmann & M Hedin. 2010. Repeated and time-correlated morphological convergence in cave-dwelling harvestmen (Opiliones, Laniatores) from montane western North America. Published in PLoS ONE.

Hedin M & SM Thomas. 2010. Molecular systematics of eastern North American Phalangodidae (Arachnida: Opiliones: Laniatores), demonstrating convergent morphological evolution in caves. Molecular Phylogenetics and Evolution 54, 107-121.