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MICHAEL G. SIMPSON, Emeritus Professor

Curator, SDSU Herbarium

Email msimpson "at" mail.sdsu.edu
Office

619.594.4479

Herbarium

619.594.8012

Fax

619.594.5676

Mailing Address

Dr. Michael G. Simpson
Department of Biology
San Diego State University
San Diego, CA 92182-4614
U.S.A.

Education: 
B.S. : University of Florida
M.S. : University of North Carolina, Chapel Hill
Ph.D. : Duke University

General Research Interests: 
Plant systematics, plant taxonomy, floristics.

Publications (*=current or former students)

2017
Aerne-Hains, L.* and M. G. Simpson. 201
7. Vegetative anatomy of the Haemodoraceae and its phylogenetic significance. International Journal of Plant Sciences 178: 117-156. (pdf)

Benet-Pierce, N. and M. G. Simpson. 2017. Taxonomic recovery of the species in the Chenopodium neomexicanum complex and description of Chenopodium sonorense sp. nov. Journal of the Torrey Botanical Society
Torrey Botanical Society 144: 339-356. (pdf)

Simpson, M. G., C. M. Guilliams*, K. E. Hasenstab-Lehman*, M. E. Mabry*, and L. Ripma*. In Press. Phylogeny of the popcorn flowers: Use of genome skimming to evaluate monophyly and interrelationships in subtribe Amsinckiinae (Boraginaceae). Taxon.

Simpson, M. G. and R. B. Kelley. 2017. Cryptantha nevadensis var. rigida elevated to species with a new name, C. juniperensis. Phytotaxa
295: 227-236. (pdf)

201
6
Chacón, J., F. Luebert, H. H. Hilger, S. Ovcinnikova, F. Selvi, L. Cecchi, C. M. Guilliams*, K. Hasenstab-Lehman*, K. Sutorý,
M. G. Simpson
, and M. Weigend. 2016. The borage family (Boraginaceae s.str.): A revised infrafamilial classification based on new phylogenetic evidence, with emphasis on the placement of some enigmatic genera. Taxon 65: 523–546. (pdf)

Luebert, F., L. Cecchi, M. W. Frohlich, M. Gottschling, C. M. Guilliams*, H. H. Hilger, K. E. Hasenstab-Lehman*, J. S. Miller, M. Mittelbach, M. Nazaire, M. Nepi, D. Nocentini, D. Ober, R. G. Olmstead, F. Selvi, M. G. Simpson, K. Sutorý, B. Valdés, G. K. Walden, and M. Weigend (Boraginales Working Group). 2016. Familial classification of the Boraginales. Taxon 65: 50
2-522. (pdf)

Mabry, M. E.*, R. A. Dowdy*, L. M. Simpson, J. P. Rebman, and M. G. Simpson. 2016. Taxonomy of the winged popcorn flower: Cryptantha pterocarya (Boraginaceae). Phytotaxa 253:97-130. (pdf)

McCabe, S. W., M. W. Dodero, and M. G. Simpson. 2016. Dudleya hendrixii A New, Rare Species From Colonet Mesa, Baja California. Madroño 63: 359-365 (pdf).

Simpson, M. G.
, L. M. Simpson, and J. P. Rebman. 2016. A new, large-flowered variety of Eremocarya micrantha (Boraginaceae). Madroño 63:39-54 (pdf).

2014
Benet-Pierce, N. and M. G. Simpson. 2014. The taxonomy of Chenopodium desiccatum and C. nitens, sp. nov. Journal of the Torrey Botanical Society 141:161-172. (pdf)

Rebman, J. P. and M. G. Simpson. 2014. Checklist of the Vascular Plants of San Diego County, 5th edition. San Diego Natural History Museum Publications. (pdf)


Ripma, L.*, M. G. Simpson, and K. Hasenstab-Lehman*. 2014. Geneious! Simplified genome skimming methods for phylogenetic systematic studies: A case study in Oreocarya (Boraginaceae). Applications in Plant Sciences doi: 10.3732/apps.1400062. (pdf)

Simpson, Michael G. 2014. Review of Intermountain Flora: Vascular Plants of the Intermountain West, U.S.A. Subclass Magnoliidae– Caryophyllidae, Vol. 2A. By N. H. Holmgren, P. K. Holmgren, J. L. Reveal, and collaborators. 2012. The New York Botanical Garden Press, Bronx, New York. 742 pp. ISBN 978-0-89327- 520-4, Price $150.00, hardcover.  Madroño 61: 146-147. (pdf)


Simpson, M. G., K. Hasenstab-Lehman*, and R. B. Kelley. 2014. Typification of the name Johnstonella (Boraginaceae). Taxon 63: 930-931. (pdf)

Simpson, M. G.
, R. Dowdy*, J. P. Rebman, R. B. Kelley, and L. M. Simpson. 2014. The recognition of two species in Eremocarya (Boraginaceae): Evidence from fornix bodies, nutlets, corolla size, and biogeography. Madroño 61: 259–275. (pdf)


2013

Guilliams, C. M.*, B. A. Veno, M. G. Simpson, and R. B. Kelley. 2013. Pectocarya anisocarpa, a new species of Boraginaceae, and a revised key for the genus in western North America. Aliso 31: 1-13. (pdf)


Silveira, M.* and Simpson, M. G. 2013. Phylogenetic systematics of the mesa mints: Pogogyne (Lamiaceae). Systematic Botany 38: 782-794. (pdf)

Simpson, M. G.
, J. P. Rebman, K. E. Hasenstab-Lehman*, C. M. Guilliams*, and P. McConnell. 2013. Cryptantha wigginsii (Boraginaceae): A presumed extinct species rediscovered. Madroño 60(1): 24-34. (pdf)


Simpson, M. G. and J. P. Rebman. 2013. A new species of Cryptantha (Boraginaceae) from the Sierra de San Pedro Mártir, Baja California, Mexico. Madroño 60(1): 35-45. (pdf) Yost, J. M., M. Bontrager, S. McCabe, D. Burton*, M. G. Simpson, K. M. Kay, and M. Ritter. 2013. Phylogenetic relationships and evolution in Dudleya (Crassulaceae). Systematic Botany 38: 1096-1104 (pdf).

2012

Dodero, M. W.* and M. G. Simpson. 2012. Dudleya crassifolia (Crassulaceae), a new species from northern Baja California, Mexico. Madroño 59: 223-229. (pdf)


Hasenstab-Lehman, Kristen E.* and M. G. Simpson. 2012. Cat’s Eyes and Popcorn Flowers: Phylogenetic Systematics of the Genus Cryptantha s.l. (Boraginaceae). Systematic Botany 37: 738–757. (pdf)  NOTE: Winner of the 2014 Grady Webster Award, given by the American Society of Plant Taxonomists, for best publication during the two year period 2012-2013!


Kelley, R. B., M. G. Simpson, and K. E. Hasenstab. 2012. Cryptantha. Pp. 455-468 in B. G. Baldwin, D. H. Goldman, D. J. Keil, R. Patterson, T. J. Rosatti, and D. H. Wilken (eds.), The Jepson Manual: Vascular Plants of California. Second Edition. University of California Press, Berkeley, Los Angeles, London.

Silveira, M., M. G. Simpson, and J. D. Jokerst. 2012. Pogogyne. Pp. 850-851 in B. G. Baldwin, D. H. Goldman, D. J. Keil, R. Patterson, T. J. Rosatti, and D. H. Wilken (eds.), The Jepson Manual: Vascular Plants of California. Second Edition. University of California Press, Berkeley, Los Angeles, London.


2010
Benet-Pierce, N. and M. G. Simpson. 2010. Chenopodium littoreum (Chenopodiaceae),  a new goosefoot from dunes of south-central coastal California. Madroño 57: 64-72. (pdf)

Guilliams, C. M.*, Simpson, M. G., and J. P. Rebman. 2011. Calyptridium parryi var. martirense (Montiaceae), a new taxon endemic to the Sierra San Pedro Mártir, Baja California, Mexico. Madroño 58(4): 258-266. (pdf)


Simpson, M. G., C. M. Guilliams*, and M. Silveira*. 2010. Taxonomy of Calyptridium parryi (Montiaceae). Madroño 57: 145-160. (pdf)


Simpson, Michael G.  2010.  Plant Systematics, 2nd edition.  Elsevier-Academic Press. (web site)

2009
Pierce, Nuri and Michael G. Simpson. 2009. Polyaperturate pollen types and ratios of heteromorphism in the monocot genus Conostylis R. Br. (Haemodoraceae). Australian Systematic Botany 22: 16-30. (pdf)

Simpson, Michael G.
and Kristen E. Hasenstab*. 2009. Cryptantha of Southern California. Crossosoma 35: 1-59. (pdf)


2007
Winner, Annette and Michael G. Simpson.  2007.  A new subspecies of Pentagramma triangularis (Pteridaceae).  Madroño 54(4): 345-353. (pdf)


2006

Rebman, Jon P. and Michael G. Simpson.  2006.  Checklist of the Vascular Plants of San Diego County.  4th edition.  San Diego Natural History Museum, San Diego, California. 


Simpson, Michael G.
2006. Plant Systematics. Elsevier-Academic Press. (web site) NOTE: Winner of the Gleason Award, New York Botanical Garden, 2006


Simpson, Michael G.
and Darren H. Burton*.  2006.  Floral anatomy of the Pontederiaceae and its systematic significance.  Aliso 22: 499-519. In: Columbus, J. T., E. A. Friar, J. M. Porter, L. M. Prince, and M. G. Simpson [eds.].  Monocots:  Comparative biology and evolution.  2 vols.  Rancho Santa Ana Botanic Garden, Claremont, California, U. S. A. (pdf)


2003
Hargrove, Lori and Michael G. Simpson.  2003. Ultrastructure of heterocolpate pollen in Cryptantha (Boraginaceae).  International Journal of Plant Sciences 164(1): 137-151. (pdf)


2000
Evans, Timothy M., Robert B. Faden, Michael G. Simpson, and Kenneth J. Sytsma.  2000.  Phylogenetic relationships in the Commelinaceae: I. A cladistic analysis of morphological data.  Systematic Botany 25: 668-691. (pdf)


1999
Marsden, Kim L.* and Michael G. Simpson.  1999.  Eryngium pendletonensis (Apiaceae), a new species from southern california.  Madroño 46(1): 55-58. (pdf)


1998
Simpson, Michael G.
  1998.  Reversal in ovary position from inferior to superior in the Haemodoraceae: evidence from floral ontogeny.  International Journal of Plant Sciences 159(3): 466-479. (pdf)


Simpson, Michael G.  1998.  Haemodoraceae.  In: Kubitski, ed. Families and Genera of Flowering Plants.   Volume IV. Flowering Plants Monocotyledons. Alismatanae and Commelinanae (except Gramineae). Pp. 212-222. Springer, Berlin. (pdf)

Simpson, Michael G. and Rudall, P.  1998.  Tecophilaeaceae.  In: Kubitski, ed. Families and Genera of Flowering Plants.   Volume III. Flowering Plants Monocotyledons. Lilianae (except Orchidaceae). Pp. 429-436. Springer, Berlin. (pdf)

1994
Simpson, Michael G.
  and Levin, G. A.  1994.  Pollen ultrastructure of the biovulate Euphorbiaceae.  International Journal of Plant Sciences 155: 313-341. (pdf)


Levin, Geoffrey A., and Michael G. Simpson.  1994.  Phylogenetic implications of pollen ultrastructure in the Oldfieldioideae (Euphorbiaceae).  Annals of the Missouri Botanical Garden 81: 203-238. (pdf


Levin, Geoffrey A., and Michael G. Simpson.  1994.  Phylogenetic relationships of Hymenocardia and Didymocistus (Euphorbiaceae).  Annals of the Missouri Botanical Garden 81: 239-244. (pdf

1993
Simpson, Michael G.
  1993.  Septal nectary anatomy and phylogeny in the Haemodoraceae.  Systematic Botany 18: 593-613. (pdf)


199
0
Simpson, Michael G.
  1990.  Phylogeny and classification of the Haemodoraceae.  Annals of the Missouri Botanical Garden 77: 722-784. (pdf)


1989
Simpson, Michael G.
  1989.  Pollen wall development of Xiphidium coeruleum (Haemodoraceae) and its systematic implications.  Annals of Botany 64: 257-269. (pdf)


1988
Simpson, Michael G.
  1988.  A critique of "Bromeliales, related monocots, and resolution of relationships among Bromeliaceae subfamilies."  Systematic Botany 13: 610-614. (pdf


Simpson, Michael G.
  1988.  Embryological development of Lachnanthes caroliniana (Haemodoraceae).  Amer. J. Bot. 75: 1394-1408. (pdf)


1987
Simpson, Michael G.
  1987.  Pollen ultrastructure of the Pontederiaceae:  evidence for exine homology with the Haemodoraceae.  Grana 26: 113-126. (pdf)


1985
Simpson, Michael G. 
1985.  Pollen ultrastructure of the Tecophilaeaceae.  Grana 24: 77-92. (pdf)


Simpson, Michael G. 
1985.  Pollen ultrastructure of the Philydraceae.  Grana 24: 23-31. (pdf)


1983
Simpson, Michael G. 
1983.  Pollen ultrastructure of the Haemodoraceae and its taxonomic significance.  Grana 22: 79-103. (pdf)


1981
Simpson, Michael G.
and W. C. Dickison.  1981.  Comparative anatomy of Lachnanthes and Lophiola (Haemodoraceae).  Flora 171: 95-113. (pdf)


Non-referred publications


Simpson, Michael G., E. Gergus, and S. C. McMillan.  2005.  Laboratory Manual for Organismal Biology: an Evolutionary Approach. 7th edition.  Rhynchops Press, La Mesa, California.

Simpson, Michael G.
  2013.  Plant Collecting and Documentation Field Notebook.  Rhynchops Press, La Mesa, California.