Oreocarya Taxonomy & Images

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Oreocarya Greene
Pittonia 1: 57. 1889


KEY TO OREOCARYA (modified from Higgins 1971)


The approximately 63 described species and 72 taxa of North American Oreocarya
(an additional 6 species/taxa (5 sp. nov. and 1 comb. nov.) in progress; see below)
[NOTE: Scale bars of all photographs = 1 mm]


ALPHABETICAL LISTING:
[Cryptantha synonyms in brackets; red=synonyms with different epithet]

Oreocarya aperta Eastwood
Oreocarya atwoodii (L. C. Higgins) Hasenstab & M.G. Simpson
Oreocarya bakeri Greene
Oreocarya barnebyi (I. M. Johnston) Hasenstab & M.G. Simpson
Oreocarya breviflora Osterhout
Oreocarya caespitosa A. Nelson
Oreocarya cana A. Nelson
Oreocarya capitata Eastwood
Oreocarya compacta (L. C. Higgins) R. B. Kelley
Oreocarya confertiflora Greene
Oreocarya crassipes (I. M. Johnston) Hasenstab & M. G. Simpson
Oreocarya creutzfeldtii (S.L. Welsh) R. B. Kelley
Oreocarya crymophila (I. M. Johnston) Jeps. & Hoover
Oreocarya depressa (M.E. Jones) J. F. Macbride
Oreocarya elata Eastwood
Oreocarya flava A. Nelson
Oreocarya flavoculata A. Nelson
Oreocarya fulvocanescens (S. Watson) Greene var. fulvocanescens
     var. nitida (Greene) R. B. Kelley
Oreocarya glomerata (Pursh) Greene
Oreocarya grahamii (I. M. Johnston) R. B. Kelley
Oreocarya hoffmannii (I. M. Johnston) Abrams
Oreocarya humilis Greene subsp. humilis
     subsp. nana (Eastwood) W. A. Weber
Oreocarya hypsophila (I. M. Johnston) Hasenstab & M.G. Simpson
Oreocarya insolita J. F. Macbride
Oreocarya interrupta Greene
Oreocarya johnstonii (L. C. Higgins) Hasenstab & M.G. Simpson
Oreocarya jonesiana Payson
Oreocarya leucophaea (Douglas) Greene
Oreocarya longiflora A. Nelson
Oreocarya mensana (M. E. Jones) Payson
Oreocarya nubigena Greene
Oreocarya oblata (M. E. Jones) J. F. Macbride
Oreocarya ochroleuca (L. C. Higgins) R. B. Kelley
Oreocarya osterhoutii Payson
Oreocarya palmeri (A. Gray) Greene
Oreocarya paradoxa A. Nelson
Oreocarya paysonii J. F. Macbride
Oreocarya propria A. Nelson & J. F. Macbride
Oreocarya revealii W. A. Weber & R. C. Wittmann
Oreocarya rollinsii (I. M. Johnston) W. A. Weber
Oreocarya roosiorum (Munz) R. B. Kelley, Hasenstab, & M.G. Simpson
Oreocarya rugulosa Payson
Oreocarya salmonensis A. Nelson & J. F. Macbride
Oreocarya schoolcraftii (Tiehm) R. B. Kelley
Oreocarya semiglabra (Barneby) Hasenstab & M.G. Simpson
Oreocarya sericea (A. Gray) Greene
Oreocarya setosissima (A. Gray) Greene
Oreocarya shackletteana (L. C. Higgins) R. B. Kelley
Oreocarya sobolifera (Payson) R. B. Kelley
Oreocarya spiculifera Piper
Oreocarya stricta Osterhout
Oreocarya subcapitata (Dorn & Lichvar) R. B. Kelley
Oreocarya subretusa (I. M. Johnston) Abrams
Oreocarya suffruticosa (Torrey) Greene var. suffruticosa
   var. abortiva (Greene) J. F. Macbride
   var. arenicola (L. C. Higgins & S. L. Welsh) R. B. Kelley
   var. cinerea (Greene) Payson
   var. laxa (J. F. Macbride) R. B. Kelley
   var. multicaulis (Torrey) Payson
   var. pustulosa (Rydberg) R. B. Kelley
   var. setosa (M. E. Jones) R. B. Kelley
Oreocarya tenuis Eastwood

Oreocarya thompsonii (I. M. Johnston) Abrams
Oreocarya thyrsiflora Greene
Oreocarya tumulosa Payson
Oreocarya virgata (Porter) Greene
Oreocarya virginensis (M. E. Jones) J. F. Macbride
Oreocarya weberi (I. M. Johnston) W. A. Weber
Oreocarya welshii (K. H. Thorne & L. C.Higgins) R. B. Kelley
Oreocarya wetherillii Eastwood

[Cryptantha aperta (Eastwood) Payson]
[Cryptantha atwoodii L.C. Higgins]
[Cryptantha bakeri (Greene) Payson]
[Cryptantha barnebyi I. M. Johnston]
[Cryptantha breviflora (Osterh.) Payson]
[Cryptantha caespitosa (A. Nelson) Payson]
[Cryptantha cana (A. Nelson) Payson]
[Cryptantha capitata (Eastwood) I. M. Johnston]
[Cryptantha compacta L.C. Higgins]
[Cryptantha confertiflora (Greene) Payson]
[Cryptantha crassipes I. M. Johnston]
[Cryptantha creutzfeldtii S. L. Welsh]
[Cryptantha crymophila I. M. Johnston]
[Cryptantha abata I. M. Johnston]
[Cryptantha elata (Eastwood) Payson]
[Cryptantha flava (A. Nelson) Payson]
[Cryptantha flavoculata (A. Nelson) Payson]
[Cryptantha fulvocanescens (S. Wats.) Payson]
     [C. f. var. nitida (Greene) R.C. Sivinski]
[Cryptantha celosioides (Eastwood) Payson]
[Cryptantha grahamii I. M. Johnston]
[Cryptantha hoffmannii I. M. Johnston]
[Cryptantha humilis (A. Gray) Payson]
     [C. humilis Payson var. nana (Eastwood) L.C. Higgins]
[Cryptantha hypsophila I. M. Johnston]
[Cryptantha insolita (J. F. Macbride) Payson]
[Cryptantha interrupta (Greene) Payson]
[Cryptantha johnstonii L.C. Higgins]
[Cryptantha jonesiana (Payson) Payson]
[Cryptantha leucophaea (Douglas) Payson]
[Cryptantha longiflora (A. Nelson) Payson]
[Cryptantha mensana (M.E. Jones) Payson]
[Cryptantha nubigena (Greene) Payson]
[Cryptantha oblata (M.E. Jones) Payson]
[Cryptantha ochroleuca L.C. Higgins]
[Cryptantha osterhoutii (Payson) Payson]
[Cryptantha palmeri (A. Gray) Payson]
[Cryptantha paradoxa (A. Nelson) Payson]
[Cryptantha paysonii (J. F. Macbride) I. M. Johnston]
[Cryptantha propria (A. Nelson & J. F. Macbride) Payson]
[Cryptantha gypsophila Reveal & C.R. Broome]
[Cryptantha rollinsii I. M. Johnston]
[Cryptantha roosiorum Munz]
[Cryptantha rugulosa (Payson) Payson]
[Cryptantha salmonensis (A. Nelson & J. F. Macbride) Payson]
[Cryptantha schoolcraftii Tiehm]
[Cryptantha semiglabra Barneby]
[Cryptantha sericea (A. Gray) Payson]
[Cryptantha setosissima (A. Gray) Payson]
[Cryptantha shackletteana L.C. Higgins]
[Cryptantha sobolifera Payson]
[Cryptantha spiculifera (Piper) Payson]
[Cryptantha stricta (Osterh.) Payson]
[Cryptantha subcapitata Dorn & Lichvar]
[Cryptantha subretusa I. M. Johnston]
[Cryptantha cinerea (Greene) Cronquist]
     [C. c. var. abortiva (Greene) Cronquist]
     [C. c. var. arenicola L.C. Higgins & S. L. Welsh]
     [C. c. var. cinerea (Greene) Cronquist]
     [C. c. var. laxa (MacBride) L.C. Higgins]
     [C. jamesii Payson var. multicaulis (Torr.) Payson]
     [C. c. var. pustulosa (Rydberg) L.C. Higgins]
     [C. jamesii Payson var. setosa (M.E. Jones) Shinners]
[Cryptantha tenuis (Eastwood) Payson]
[Cryptantha thompsonii I. M. Johnston]
[Cryptantha thrysiflora (Greene) Payson]
[Cryptantha tumulosa (Payson) Payson]
[Cryptantha virgata (Porter) Payson]
[Cryptantha virginensis (M.E. Jones) Payson]
[Cryptantha weberi I. M. Johnston]
[Cryptantha welshii K.H.Thorne & L.C.Higgins]
[Cryptantha wetherillii (Eastwood) Payson]

IN PROGRESS:  

Oreocarya coryi (I. M. Johnston) R. B. Kelley & Ripma, comb. nov.
Oreocarya flumenavia
R. B. Kelley & Ripma, sp. nov.
Oreocarya higginsii R. B. Kelley & Ripma, sp. nov.
Oreocarya howelliana R. B. Kelley & Ripma, sp. nov.
Oreocarya montana R. B. Kelley & Ripma, sp. nov.
Oreocarya ursina R. B. Kelley & Ripma, sp. nov.




INFORMAL GROUPS, after Higgins 1971 (=Higgins, L. C. 1974. Southw. Naturalist 19: 127; **=not treated by Higgins)
[Blue=Corolla>calyx, heterostylous, nutlets smooth; Purple=Corolla>calyx, heterostylous, nutlets rough]

The jamesii group(nutlets hemispheric, smooth or rugose/tuberculate):
     Oreocarya palmeri (A. Gray) Greene  
     Oreocarya suffruticosa (Torr.) Greene [& vars.]
     Oreocarya crassipes (I. M. Johnston) Hasenstab & M.G. Simpson  
     Oreocarya oblata (M.E. Jones) J. F. Macbride  
     Oreocarya paysonii J. F. Macbride  

The flava group (nutlets compressed, scar closed):
     Oreocarya atwoodii (L.C. Higgins) Hasenstab & M.G. Simpson†  
     Oreocarya capitata Eastwood  
     Oreocarya confertiflora Greene  
     Oreocarya flava A. Nelson  
     Oreocarya johnstonii (L.C. Higgins) Hasenstab & M.G. Simpson  
     Oreocarya leucophaea (Douglas) Greene  
     Oreocarya semiglabra (Barneby) Hasenstab & M.G. Simpson  
     Oreocarya salmonensis A. Nelson & J. F. Macbride  

The stricta group(stout, hispid stems; nutlet ventral surface smooth):
     Oreocarya barnebyi (I. M. Johnston) Hasenstab & M.G. Simpson  
     Oreocarya stricta Osterh.  

The nubigena group (nutlets lance-ovate, ventrally smooth, dorsally rough):
 
    Oreocarya crymophila (I. M. Johnston) Jeps. & Hoover  
     Oreocarya howelliana R.B. Kelley, sp. nov.**
     Oreocarya hypsophila (I. M. Johnston) Hasenstab & M.G. Simpson  
     Oreocarya nubigena Greene  
     Oreocarya schoolcraftii (Tiehm) R.B. Kelley  
     Oreocarya subretusa (I. M. Johnston) Abrams  
     Oreocarya ursina R.B. Kelley, sp. nov.**
     
The abata group(nutlets rugose or tuberculate; ventral scar open, triangular, with elevated margin):
     Oreocarya depressa (M.E. Jones) J. F. Macbride  
     Oreocarya hoffmannii (I. M. Johnston) Abrams  
     Oreocarya tumulosa Payson  
     Oreocarya virginensis (M.E. Jones) J. F. Macbride  

The caespitosa group(hummock forming vs. gypsophilous):
     Oreocarya caespitosa A. Nelson  
     Oreocarya ochroleuca (L.C. Higgins) R.B. Kelley  

The humilis group (nutlets muricate, scar open, basally triangular):
     Oreocarya breviflora Osterh.  
     Oreocarya cana A. Nelson  
     Oreocarya fulvocanescens (S. Wats.) Greene [& vars.]
     Oreocarya jonesiana Payson  
     Oreocarya compacta (L.C. Higgins) R. B. Kelley  
     Oreocarya humilis (A. Gray) Greene [& subspp.]
     Oreocarya propria A. Nelson & J. F. Macbride  
     Oreocarya roosiorum (Munz) R.B. Kelley, Hasenstab, & M.G. Simpson  

The elata group(corolla tube < calyx [except O. rollinsii] nutlets dorsally roughened, rugose and/or tuberculate/muricate, ventrally smooth or rough; scar straight, margins not elevated):
     Oreocarya aperta Eastwood  
     Oreocarya elata Eastwood  
     Oreocarya glomerata (Pursh) Greene
     Oreocarya grahamii (I. M. Johnston) R.B. Kelley
     Oreocarya interrupta Greene  
     Oreocarya rollinsii (I. M. Johnston) W.A. Weber  
     Oreocarya rugulosa Payson  
     Oreocarya shackletteana (L.C. Higgins) R.B. Kelley  
     Oreocarya sobolifera (Payson) R.B. Kelley  
     Oreocarya spiculifera Piper  
     Oreocarya thompsonii (I. M. Johnston) Abrams  
     Oreocarya thyrsiflora Greene  
     Oreocarya weberi (I. M. Johnston) W.A. Weber  

The flavoculata group(nutlets rugose and tuberculate or muricate or foveolate; scar slightly open [except O. bakeri], constricted above base, margin usu. elevated):
     Oreocarya bakeri Greene  
     Oreocarya flavoculata A. Nelson  
     Oreocarya longiflora A. Nelson  
     Oreocarya mensana (M. E. Jones) Payson  
     Oreocarya osterhoutii Payson  
     Oreocarya paradoxa A. Nelson  
     Oreocarya tenuis Eastwood  
     Oreocarya wetherillii Eastwood  

The virgata group(stem stout, fistulose; inflorescene spicate with elongated foliar bracts):
     Oreocarya virgata (Porter) Greene  

The setosissima group(stem simple, erect; nutlets broadly winged; calyx segments ovate):
     Oreocarya setosissima (A. Gray) Greene  




Unplaced to date:
     Oreocarya creutzfeldtii (S.L. Welsh) R. B. Kelley  
     Oreocarya revealii W.A. Weber & R.C. Wittmann  
     Oreocarya sericea Greene  [treated by Higgins 1971, but not placed in a group]
     Oreocarya subcapitata (Dorn & Lichvar) R.B. Kelley  
     Oreocarya welshii (K.H.Thorne & L.C.Higgins) R.B. Kelley