Andersonglossum Cryptantha Eremocarya Greeneocharis Harpagonella Pectocarya Plagiobothrys Simpsonanthus
Adelinia Amsinckia Amsinckiopsis Andersonglossum Cryptantha Dasynotus Eremocarya Greeneocharis Gruvelia Harpagonella Johnstonella Oncaglossum Oreocarya Pectocarya Plagiobothrys Simpsonanthus Sonnea

Systematics of Amsinckiinae (Boraginaceae): The Popcorn Flowers

[Amsinckiinae HOME]

Greeneocharis Gürke & Harms

Nat. Pflanzenfam. [Engler & Prantl] Regist. 460. 1899

after California botanist Edward Lee Greene (1843–1915) + charis, "beauty, delight"

Piptocalyx Torrey, nom. illegit. (later homonym), U.S. Expl. Exped., Phan. Pacific N. Amer. 17(2): 414, t. 12. 1874; C. Wilkes Expl. Exped.
Wheelerella G.B.Grant, nom. illegit. (superfulous name), Bull. S. Calif. Acad. Sci. 5: 28. 1906

Type Species/EspeciesTipo:
Greeneocharis circumscissa (Hooker & Arnott) Rydberg, Bull. Torrey Bot. Club 36: 677. 1909

Distribution of Greeneocharis:
LEFT: POWO. 2019. Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet. Accessed 26 Dec 2019.
RIGHT: GBIF. 2019. Global Biodiversity Information Facility. Accessed 26 Dec 2019.

2 species and 3 minimum ranked taxa
(NA=North America; SA=South America)

Greeneocharis circumscissa (Hooker & Arnott) Rydberg  NA & SA
     G. c. var. circumscissa   NA & SA
     G. c. var. rosulata (J.T.Howell) Hasenstab & M.G.Simpson  NA
Greeneocharia similis (K.Mathew & P.H.Raven) Hasenstab & M.G.Simpson  NA
[Cryptantha circumscissa (Hooker & Arnott) I.M.Johnston]
[Cryptantha circumscissa (Hooker & Arnott) I.M.Johnston var. circumscissa]
[Cryptantha circumscissa (Hooker & Arnott) I.M.Johnston var. rosulata J.T.Howell]
[Cryptantha similis K.Mathew & P.H.Raven]

California Native Plant Society (CNPS) Inventory Listings:

1B = Rare, threatened, or endangered in California and elsewhere
.2 = Fairly endangered in California (20-80% occurrences threatened)

KEY TO TAXA (after Kelley et al. 2012)
Corolla limb 3.5–6 mm diam, appendages larger than small to minute, yellow ............................................................................. G. similis
Corolla limb generally 1–2 mm diam, appendages small to minute, white to pale yellow .................................................... G. circumscissa
      Stem generally > 2 cm; calyx 3–4 mm in fruit; fruit axis ± = mature nutlets; nutlet beak 0; leaf margin rolled under ..... var. circumscissa
      Stem 0.5–2 cm; calyx 2.5–3 mm in fruit; fruit axis < mature nutlets; nutlet strongly beaked; leaf margin ± flat .................... var. rosulata

NOTE: Greeneocharis similis and G. circumscissa also differ in chromosome number and pollen diameter, major reasons for their status as separate species. The chromosome number of Greeneocharis circumscissa var. rosulata has not been determined, but pollen diameter is reported to be similar to G. circumscissa var. circumscissa. (See Mathew & Raven 1962).
Greeneocharis similis n=6 pollen 5.5-6.5 µm in diameter
Greeneocharis circumscissa var. circumscissa n=12 (18) pollen 7-9 µm in diameter



Gürke, R. L. A. M. and H. A. T. Harms. 1899. Greeneocharis. Pp. 460 in H. G. A. Engler and K. A. E. Prantl (eds.), Naturlichen Pflanzenfamilien Registr. W. Engelmann, Leipzig.
Kelley, R. B., M. G. Simpson, and K. E. Hasenstab. 2012. Cryptantha. Pp. 455-468 in B. G. Baldwin, et al. (eds.), The Jepson Manual: Vascular Plants of California. Second Edition.
      University of California Press, Berkeley, Los Angeles, London.

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